how to Present the special-temporal distribution CHART of all hominids (7mya to present)
Hominid
refers to any of the modern or extinct bipedal primates of the family hominidae including all species of the
general Homo and Australopithecus. In biology and ecology, extinction is the end of
an organism or of a group of organisms (taxon), normally a species. (World
English Dictionary)
Generally
hominid refers to any human like species including a modern man. They are
biological family to which human being belong. Our immediate evolutionary
family is comprised of the hominoids, the group of primates that includes the
"lesser apes" (siamangs and
gibbons) as well as the "great
apes" (chimpanzees, bonobos, gorillas, and orangutans).
The ancestors of hominids are regarded as great Apes including chimpanzee, bonobos, gorillas and orangutans, among the great ape our closest
relatives are chimpanzees, and bonobos. The fossil record, along with
studies of human and ape DNA, indicate that humans shared a common ancestor
with chimpanzees and bonobos sometime around 6 million years ago (mya). However
this does not means that the modern man was evolved from the chimpanzees but we
just share the common ancestor.
Hominids
are linked to theory of evolution of man propounded by Charles Darwin
(1809-1882), he claimed that a great variety of plant and animal life in the
earth are developed gradually through natural processes, but species change or
adopt over time in response to the environment. His theory of “Natural
Selection” based on observation that members of single species vary greatly in
shape size, color, strength to which he believed that these variations are
inherited. However fossil records along with the study of Human and Apes DNA
indicate that human being is similar to Great Apes family especially champs in
genetic make up about 99 %. Before 7mya they shared the same ancestor but after
this era they diverged.
Hominids
especially human are considered to be best and most intelligent biological
family on the earth or planet regarding the cranium capacity of the brain they
possess. The homo genus are distinguished from apes because of bipedalism
(walking and standing with two limbs) and using simple tools such as sticks to
catch the insects.
The
hominin lineage origin can be understood from the ancestral state, for our
lineage The Human-Chimpanzee Last Common Ancestor (HC-LCA) is the species from
which the hominin lineage and the chimpanzee & bonobo lineage diverged.
This was during the Miocene about 7 million years ago (7mya), where there was
great diversity of ape species with dozen of species known from the fossil
record across Africa, Europe and Asia. These
species varied in their anatomy and ecology, and it is not clear which, if any,
of the fossil species discovered thus far represent the Human-Chimpanzee Last Common
Ancestor, although we know from fossil and comparative evidence that it was
much more similar to living apes than to living humans. (Kunimatsu et al. 2007;
Young and MacLatchy, 2004)
The
Humman-Chempanzee Last Common Ancestror would have had an ape-sized brain and
body, with relatively long arms and fingers and a grasping foot that allowed it
to forage in the trees. The canine teeth were probably large and sharp, as seen
in several Miocene hominoids. Moreover, the canines were probably sexually
dimorphic, with males having much larger canines than females, as seen among
the living great apes and Miocene fossils. Like living apes it would have
walked quadrupedally (on all fours) when on the ground, and its diet would have
consisted almost entirely of plant foods, primarily fruit and leaves.
The
following is the distribution of the hominids from seven millions years ago to
present (7mya to present); -
Sahelanthropus tchadensis
from Chad and Orrorin tegenensis from
Kenya are the oldest hominins currently known. In both Orrorin and Sahelanthropus
the canine teeth of males are larger and more pointed than in modern humans,
but are small and blunt compared to the canines of male apes. This suggests that
canine sexual dimorphism and by extension, competition among males for mating
access to females was diminished in these early hominins compared to the great
apes.
Sahelanthropus
tchadensis, named TM 266 (Toumai) discovered in 2001 Chad by Alain
Beauvilain, Fanone Gongdibe, Mahamat Adoum and Ahounta Djimdoumalbaye and dated
to 7 mya, is known from a largely complete skull and some other fragmentary
remains. Its brain size, 360cc, is within the range seen in chimpanzees, and
the skull has a massive brow ridge, similar in thickness to male gorillas. However,
the position and orientation of the foramen magnum, the hole in the base of the
skull through which the spinal cord passes, suggests that Sahelanthropus stood and walked bipedally, with its spinal column
held vertically as in modern humans rather than horizontally as in apes and
other quadrupeds.
Orrorin
tugenensis, named Bar 1000’00 discovered in 2000 in Kenya by Martin
Pickford, Kiptalam Cheboi, Dominique Gommery, Pierre Mein, and Brigitte Senut
and dated to about 6 mya, is known primarily from postcranial fossils,
including a partial femur. The proximal portion of the femur shows similarities
to those of modern humans, suggesting the species was bipedal. No skulls of Orrorin have been recovered, and so it’s
cranial morphology and brain size are uncertain.
Ardipithecus ramidus,
named Ardi from Ethiopia discovered in 1994 by Yohannes Haile Selassie
and dated to 4.4 mya, is known from a nearly complete skeleton as well as
numerous other dental and skeletal remains. Ardipithecus
ramidus and an older related species known from fragmentary remains Ardipithecus kadabba (5.8–5.2 mya), have
reduced canines similar to those of Orrorin
and Sahelanthropus. The skull of Ardipithecus ramidus is rather ape-like
and broadly similar to that of Sahelanthropus,
with a small chimpanzee-sized brain of 300–350cc. The Ardipithecus postcranial skeleton is intriguing. Although badly
fragmented, the pelvis recovered reveals morphology quite different from that
of living apes, with a shorter, more bowl-like shape that strongly suggests Ardipithecus walked bipedally; this is
consistent with the foramen magnum position, which suggests an upright posture.
However, its long forelimbs and fingers and its divergent, grasping first toe
(hallux) suggest Ardipithecus spent
much of its time in the trees. The overall impression is of a largely arboreal
species that walked bipedally whenever it ventured to the ground.
Australopithecus,
were discovered around 4mya they were hominins which were adept terrestrial
bipeds but continued to use the trees for food and protection. The first
specimens of Australopithecus were
discovered in South Africa in 1924 and named Taung 1 (Taung Child) such
as Australopithecus africanus
discovered by Raymond Dart dated 2.5 mya. (Dart, 1925), and research efforts
over the subsequent eight decades have produced hundreds of fossils from
several species at sites all across East and Southern Africa. Australopithecus was a highly successful
genus that persisted for nearly three million years. The best-known Australopithecus species are Australopithecus afarensis (3.6–2.9 mya)
like Australopithecus afarensis named
LH 4 discovered in 1974 at Laetoli, Tanzania by Donald Johanson dated
about (2.9 – 3.9 mya) from East Africa and Australopithecus
africanus (3.2–2.0mya) from South Africa. The pelvis and lower limb of
these species clearly indicates that they were fully bipedal: the pelvis is
short and bowl-shaped, bringing the gluteal muscles around to the side of the
body, as in modern humans, for trunk stabilization during bipedalism, and the
first toe is in line with the other toes (Ward, 2002; Harcourt-Smith and
Aiello, 2004).
The Australopithecus
foot may even have had a human-like arch, based on analysis of the metatarsals
and the fossilized Laetoli footprints (Ward et al. 2011). the forearms were
long and the fingers and toes were long and somewhat curved, suggesting that Australopithecus regularly used the
trees to forage and perhaps as a refuge from predators at night. This mixed
terrestrial & arboreal strategy would have served these species well in the
mixed woodland and savannah environments they inhabited. Brain size in Australopithecus ranged between 390 and
515cc, similar to chimpanzees and gorillas (Falk et al. 2000), suggesting
cognitive abilities were broadly similar to living apes. Body size in Australopithecus was rather small and
sexually dimorphic, about 30kg for females and 40kg for males (McHenry, 1992).
This level of dimorphism is not reflected in the canines, which were small,
blunt, and monomorphic as in earlier hominins.
Paranthropus robust, is
a subgroup of Australopithecus, they
are named as separate genus because of their enormous teeth and chewing
muscles, took this adaptation to the extreme. Most Australopithecus species were extinct by 2 mya, but some robust forms persisted until about 1.2
mya in East and South Africa. For instance Paranthropus
robusts named TM 1517 discovered 1938 in South Africa by Gert
Terblanche dated about 2 mya.
Homo,
these were the earliest genus of our own, they are early specimens similar in
brain and body size to Australopithecus,
but show differences in their molar teeth, suggesting a change in diet. Indeed,
by at least 1.8 mya, early members of our genus were using primitive stone
tools to butcher animal carcasses, adding energy-rich meat and bone marrow to
their plant-based diet.
The
oldest member of the genus Homo, is Homo habilis dated (2.3–1.4 mya) is
found in East Africa and is associated with butchered animal bones and simple
stone tools, like Homo rudolfensis named Uraha 501,
discovered in 1991 in Malawi by Tyson Msiska, Timothy Bromage and Friedemann
Schrenk.
Homo erectus
is more formidable and widespread descendant found throughout Africa and
Eurasia and persisted from 1.9 mya to 100 kya, and perhaps even later (Anton,
2003). Like modern humans, Homo erectus lacked
the forelimb adaptations for climbing seen in Australopithecus. Its global expansion suggests Homo erectus was ecologically flexible,
with the cognitive capacity to adapt and thrive in vastly different
environments. Not surprisingly, it is with Homo
erectus that we begin to see a major increase in brain size, up to 1,250cc
for later Asian specimens (Anton, 2003). Molar size is reduced in Homo erectus relative to Australopithecus, reflecting its softer,
richer diet. For instance Homo erectus
named D2700 dicovered in 2001 Georgia dated about 1.8 mya. Also Homo erectus named Yuanmou Man
discovered in 1965 China by Fang Qian dated about 1.7 mya to between 0.5 and
0.6 mya. Homo erectus named Sangiran
2 discovered in 1937 Indonesia by G. H. R. von Koenigswald dated about 0.7
– 1.6 mya.
Homo heidelbergensis,
is a species very much like us in terms of body proportions, dental
adaptations, and cognitive ability followed Homo
erectus (Rightmire, 2009). Homo
heidelbergensis, often referred to as an "archaic" Homo sapiens,
was an active big-game hunter, produced sophisticated Levallois style tools,
and by at least 400 kya had learned to control fire (Roebroeks and Villa, 2011).
For instance Homo heidelbergensis or
Homo erectus named Madam Buya discovered in 1997 Eritrea by Ernesto
Abbate dated about 600k – 1.4 mya, another was Homo heidelbergensis named Bodo discovered in 1976 Ethiopia
by A. Asfaw dated 600 kya. As well as Homo
heidelbergensis named Mauer 1 (Heidelberg Man) discovered in 1907 Germany
by Daniel Hartmann dated 500 kya.
Homo neanderthalensis,
evolved from Homo heidelbergensis
populations in Europe by at least 250 kya, they are said to be cold-adapted
hominins with stout physiques, complex behaviors, and brains similar in size to
ours, for instance Homo neanderthalensis named
Altamura Man discovered in Italy dated 250 kya. Homo neanderthalensis named Bontnewydd(Pontynewydd) discovere
in 1981 United Kingdom dated 230 kya.
Homo sapiens
in which Fossil and DNA evidence suggests that it’s our own species evolved in
Africa 200 kya, probably from Homo
heidelbergensis. The increased behavioral sophistication of Homo sapiens, as indicated by our large
brains (1,400cc) and archeological evidence of a broader tool set and clever
hunting techniques, allowed our species to flourish and grow on the African
continent. For instance Homo sapiens
named Omo 1 discovered in 1967 Ethiopia by Richard Leakey dated 190 kya.
Also Homo sapiens named Jebel Irhoud
1 discovered in 1991 Morocco dated 160 kya.
Homo neanderthals,
were displaced from our species and they were in Europe and Asia. By 100kya,
our species spilled into Eurasia, eventually expanding across the entire globe
into Australia and the Americas. Studies of ancient DNA extracted from Neanderthal fossils suggest our species
may have occasionally interbred with them. For instance Homo naenderthalensis named Krapina discovered in 1899
Croatia by Dragutin Gorjanovic Kramberger dated 100 kya – 127 kya. Homo naenderthalensis named Scladina
discovered in Belgium dated 80 kya – 127 kya. Also Homo naenderthalensis named La Ferrassie 1 discovered in
1909 in France by R. Capitan and D. Peyrony dated 70 kya.
The
following is the CHART representing the distribution of the hominids as
explained above.
NAME
|
AGE
|
SPECIES
|
YEAR DESCOVERED
|
COUNTRY
|
DISCOVERED BY
|
TM 266(Toumai)
|
7 mya
|
Sahelanthropus
tchadensis
|
2001
|
Chad
|
Alain Beauvilain, Fanone Gongdibe,
Mahamat Adoum and Ahounta Djimdoumalbaye
|
BAR 1000’00
|
6 mya
|
Orrorin
tugenensis
|
2000
|
Kenya
|
Martin Pickford, Kiptalam Cheboi,
Dominique Gommery, Pierre Mein, Brigitte Senut
|
Ardi
|
4.4 mya
|
Ardipithecus
ramidus
|
1994
|
Ethiopia
|
Yohannes Haile Selassie
|
Taung 1 (Taung Child)
|
2.5 mya
|
Australopithecus
africanus
|
1924
|
South Africa
|
Raymond Dart
|
LH 4
|
2.9 3.9 mya
|
Australopithecus
afarensis
|
1974
|
Laetoli, Tanzania
|
Donald Johnson
|
TM 1517
|
2 mya
|
Paranthropus
robustus
|
1938
|
South Africa
|
Gert Terblanche
|
Uraha 501
|
2.3-2.5 mya
|
Homo
rudolfensis
|
1991
|
Malawi
|
Tyson Msiska, Timothy Bromage, Friedemann
Schrenk
|
D2700
|
1.8 mya
|
Homo
erectus
|
2001
|
Georgia
|
|
Yuanmou Man
|
1.7 mya/0.5-0.6mya
|
Homo
erectus
|
1965
|
China
|
Fang Quian
|
Sangiran 2
|
0.7-1.6 mya
|
Homo
erectus
|
1937
|
Indonesian
|
G. H. R. von Koenignswald
|
Madam Buya
|
600k-1.4mya
|
Homo
heidelbergensis or Homo erectus
|
1997
|
Eritrea
|
Erasto Abbate
|
Bodo
|
600k
|
Homo
heidelbergensis or Homo erectus
|
1976
|
Ethiopia
|
A.
Asfaw
|
Mauer 1 (Heidelberg Man)
|
500k
|
Homo
heidelbergensis
|
1907
|
Germany
|
Daniel Hartmann
|
Altamura Man
|
250k
|
Homo
neanderthalensis
|
|
Italy
|
|
Bontnewydd
(Pontynewydd)
|
230kya
|
Homo
neanderthalensis
|
1981
|
UK
|
|
Omo 1
|
190kya
|
Homo
sapiens
|
1967
|
Ethiopia
|
Richard Leakey
|
Jebel Irhoud1
|
160kya
|
Homo
sapiens
|
1991
|
Morocco
|
|
Krapina
|
100kya-127kya
|
Homo
neandthalensis
|
1899
|
Croatia
|
Dragutin Garjanovic-Kramberger
|
Scladina
|
80kya-127kya
|
Homo
neanderthalensis
|
|
Belgium
|
|
La Ferrassie 1
|
70kya
|
Homo
neanderthalensis
|
1909
|
France
|
R. Capitan and D. Peyrony
|
Therefore,
the evolution of man from 7mya as
represented by distribution of hominids shown above had is showing how
processes of man evolution were taking place as the hominids were sharing their
ancestral and they were evolving from one stage of change to another in the aid
of the environmental changes.
REFERENCES
Anton,
S. C. (2003). “Natural history of Homo erectus”. American Journal of Physical Anthropology
S37, 126-70.
Dart,
R.A. (1925). “Australopithecus africanus”: the southern ape-man of
Africa. Nature 115, 195-199.
Falk,
D. et al. (2000). “Early hominid
brain evolution: a new look at old endocasts”. Journal of Human
Evolution 38, 695-717.
Kunimatsu,
Y. et al. (2007). “A new Late
Miocene great ape from Kenya and its implications for the origins of African
great apes and humans”. PNAS USA 104, 19661-19662.
McHenry,
H. M. (1992). “Body size and proportions
in early hominids”. American
Journal of Physical Anthropology 87, 407-431
Ward,
C. V. et al. (2011). “Complete
fourth metatarsal and arches in the foot of Australopithecus afarensis”. Science 331, 750-753
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